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Phylum
Mollusca
Class
Gastropoda
Family
Littorinidae
Periwinkles
| Periwinkles are algal grazers found high above the waterline on rocky
shores. They move vertically with the tide. They are capable
of trapping water within the shell and hibernate for extended periods if
required. |
HAWAII
Littoraria
(Littorinopsis) intermedia
Kaupo, Oahu
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Mokuleia, Oahu |
Diamond Head, Oahu |
Portlock, Oahu |
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DOTTED PERIWINKLE
| Abundant in the spray zone. Attains 1.25
inch. Indo-Pacific. Formerly known as Littorina pintado. |
Littoraria
(Littoraria) coccinea
 Portlock, Oahu |
Portlock, Oahu |
With L. intermedia, Portlock, Oahu |
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SCARLET PERIWINKLE
| Very rare in the spray zone in Hawaii but abundant throughout the
Central & Southwest Pacific. Attains 1 inch. |
Littoraria
(Littorinopsis) scabra scabra
Lilipuna Pier, Kaneohe Bay, Oahu |
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ROUGH or MANGROVE
PERIWINKLE
| Common on seawalls and mangrove roots in bays and harbors. Attains
1.5 inches. Hawaii & the Indo-Pacific. |
Nodilittorina
(Nodilittorina) hawaiensis 
Kaupo, Oahu
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Kaupo, Oahu |
Kaupo, Oahu
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Mokuleia, Oahu |
Mokuleia, Oahu
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Portlock, Oahu |
Diamond Head, Oahu |
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HAWAIIAN
PERIWINKLE
| Found on rocky shores just above the waterline. Endemic to
Hawaii. Color and sculpture is variable. Attains 1/2 inch.
Formerly known as Nodilittorina picta. |
Peasiella
tantilla
 Diamond
Head, Oahu |
Diamond Head, Oahu
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Diamond Head, Oahu
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Diamond Head, Oahu |
Puako, Hawaii |
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KEELED PERIWINKLE
| Common in small pools & crevices in the high intertidal zone.
Adults develop pronounced spiral keels. Attains 1/4 inch.
Hawaii & the Indo-Pacific. |
Family
Planaxiidae
Cluster
Winks
| Thick shells that are usually found in tight clusters on rocky shores. |
HAWAII
Planaxis
labiosa
Portlock, Oahu
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Portlock, Oahu |
Portlock, Oahu
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Portlock, Oahu |
Portlock, Oahu
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Portlock, Oahu |
Kapoho Point, Oahu |
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DWARF PACIFIC PLANAXIS
| Abundant under stones on rocky shores with fresh water seeps.
Shell covered with a fuzzy brown periostracum. Attains
1/2 inch. Hawaii through the Southwest Pacific. |
Subclass Pulmonata
Air-Breathers
Family
Melampidae
Melampid
Shells
| Melampids are air-breathing snails found along rocky shorelines,
especially where freshwater enters the sea. They do not have
gills. |
HAWAII
Allochroa
bronni
Kapoho Point, Oahu |
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| An endemic air-breathing snail found in under stones or gravel the splash
zone . Attains
1/2 inch. |
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